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Bibliography of Monitor Lizards

Extract from A Little Book of Monitor Lizards © D. Bennett 1995. Viper Press, UK

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English translations of papers marked # are available here >>

ABBASI,A. & BRAUNITZER,G. 1991. Primary structure of haemoglobin from monitor lizard (Varanus exanthematicus albigularis). Biol. Chem. Hoppe-Seyler. 372:473-479.
D'ABREU,E.A. 1933. J. Bombay. Nat. Hist. Soc. 36 (2):269-270. Notes on monitor lizards.
AHL,E. 1932. Eine neue Eidechse und zwei neue Frosche von der insel Jobi. Mitt.Zool.Mus. Berlin 17(6):892-899
AKITA,K. 1992. An anatomical investigation into the muscles of the pelvic outlet in Iguanas (Iguana iguana) and Varanus (Varanus dumerilii) with special reference to their nerve supply. Ann.Anat. 174:119-129.
AKTAN,F. 1971. Guneydogu Anadoluda Varanus griseus Daud. (col Varani). In Turkish and German with English summary. Turk.Biol.Derg. 21:106-117.
ALBERTS,A. 1994. Off to see the lizard: lessons from the wild. The Vivarium.5(5):26-28.
ALI,S. 1944. J.Bomb.Nat.Hist.Soc. 44:479-480 Courtship of the monitor lizard (Varanus monitor).
ANDERSON,J. 1872. Proc. Zool. Soc. London 26:371-404. On some Persian, Himalayan and other reptiles.
ANDERSON,J. 1895. Proc.Zool.Soc.London 635-663. On a collection of reptiles and batrachians made by Col. Yerbury at Aden and its neighbourhood.
ANDERSON,J. 1889. Report on the mammals, reptiles an batrachians, chiefly from the Mergui Archipelago, collected for the trustees of the Indian Museum. J.Linn.Soc.Zool. 21:331-350.
ANDERSON,J. 1898. Zoology of Egypt 1. Reptiles and Batrachia.Bernard Quaritch, London.
ANDERSON.C. 1931. Reconstruction of the head of Melagania prisca. Aus.Mus.Mag. 4:147-148.
ANDERSON,S.C. 1963. Amphibians and reptiles from Iran. Proc. Cali. Acad. Sci.31(16):417-498.
ANDERSON,S.C. & LEVITON,A.E. 1969  Reptiles and Amphibians of Afghanistan. Proc Calif.Acad.Sci. 37(2):25-56.
ANDRES,A. 1904. Intelligenz eine wustenwarens (Varanus griseus). Bl. Aqua. Terrar. Kunde. 15:269.
ANDREWS,H.V. 1995. Sexual maturation in Varanus salvator (LAURENTI 1768) with notes on growth and reproductive effort. Herpetological Journal 5 (1):189-194.
ANDREWS,H. & GAULKE,M. 1990. Observations of the reproductive biology and growth of the water monitor (Varanus salvator) at the Madras Crocodile Bank. Hamadryad 15(1):1-5.
ANNANDALE,N. 1921. The reptiles and batrachia of Barkuda Island. Rec.Ind.Mus. 22(4).
ANON. 1909. A bird enemy - the goanna. Emu 8:157-158.
ANON? 1937. A Guide to the Vertebrate Fauna of the Eastern Cape Province South Africa. Pt. 2 Reptiles, amphibians and freshwater fishes. Albany Museum, Grahamstown.
ANON. 1963. Aus befreundeten Zoos: (Varanus giganteus in Stuttgart Wilhelma Zoo). Freunde Kolner Zoo 6 (7):122.
ANON. 1965. Neues aus der Wilhelma: Geburt von Varanus mertensi. Aquar.Terrar. 16 (9):306-307.
ANON.1968 New Scientist 40:352. Komodo dragons breed in captivity.
ANON. 1978. Introducing the Irulas. Hamadryad 3(1):8-9.
ANON. 1980. Int.Zoo News. 27(5/6):51-52. Timor monitor lizard birth may be the first for the U.S.A.
ANON. 1981. Captive breeding of the Malayan water monitor (Varanus salvator) at San Antonio Zoo. AAZPA BEAN Award application.
ANON. 1983a. .A record size water monitor lizard (7.5 feet long) in Sabah, East Malaysia. Hamadryad 8 (3):Cover
ANON. 1983b. Traffic Bulletin 4 (6):71-79. International trade in skins of monitor and tegu lizards 1975-1980
ANON. 1984 Tempo 8(8): Food Trip:  Dressed bayawak. (Reprinted in GAULKE 1984).
ANON. 1987. V.exanthematicus microstictus in Reptilenzoo Iguana. Report from Reptilienzoo Iguana, Vlissingen, Netherlands.#
ANON. 1988. AAZPA BEAN Award application. Philadelphia Zoo. Breeding the mangrove monitor Varanus indicus.
ARCHER,M. & WADE,M. 1976. Results of the Ray M. Lenley Expedition Pt. 1. The Allingham Formation and a new Pliocene vertebrate from northern Queenland. Mem. Queensland Mus. 17 (3): 379-397.
ARMSTRONG,G. 1981. South Australian Herp. Group Newsletter 1981 (March):3-4. Northern Territory Field Trip Report: Victoria River and Mataranka areas 17-19 October 1980.
ARNOLD,E.N. 1984. Journal of Zoology (London). 204: 329-354. Ecology of lowland lizards in the eastern United Arab Emirates.
ARNOLD 1986. Fauna Saudi Arabia 8: 385-435. A key and annotated checklist to the lizards and amphisbaenians of Arabia.
ATAEV,C.A. 1987. Comparative aspects of the river valleys of the Sumbar and the Chandyr. Izv. Akad.Nauk. Turkmen SSR Ser. Biol. Nauk. 6: 49-54.  (in Russian).
ATTWELL,R.I.G. 1966. Possible bird-crocodile commensalism in Zambia. Ostrich 37:54-55.
AUERBACH,R.D. 1985. The reptiles of Gaborone. Botswana Book Centre, Gaborone.
AUERBACH,R.D. 1995. Segwagwanyana. Ein beitrage zur afrikanischer Ethnoherpetologie. Edition Chimera bei Bucher Kreth, Frankfurt Main.
AUFFENBERG,W. 1970. A day with number 19 - A report on a study of the Komodo monitor. Animal Kingdom. 73:18-23
AUFFENBERG,W. 1972. Komodo dragons. Nat.Hist. 81 (4): 52-59.
AUFFENBERG,W. 1976. First description of an adult Varanus grayi. Copeia (3):586-588).
AUFFENBERG,W. 1978a. Grays monitor lizard - status survey. World Wildlife Yearbook 1977-1978:129-131.
AUFFENBERG,W.  1978b. In GREENBERG & MACLEAN (Eds.) Behaviour and Neurology of Lizards: 301-331. Govt. Printing Office, Washington DC. U.S.A.  Social and feeding behaviour in Varanus komodoensis.
AUFFENBERG,W. 1979a . A monitor lizard in the Philippines. Ornyx  15 (1):39-46.
AUFFENBERG,W. 1979b. Intersexual differences in behaviour of captive Varanus bengalensis. J.Herpet. 13(3):313-315.
AUFFENBERG,W. 1979c. . Dectective Auffenberg gets his beast. Int.Wldf. 9(2): 13-15
AUFFENBERG,W. 1979d. Research on monitor lizards. Tiger Paper 6(4):20-21.
AUFFENBERG,W. 1980.  The herpetofauna of Komodo with notes on adjacent areas. Bull.Fla.State.Mus.Biol.Sci 25(2):39-156
AUFFENBERG,W. 1981. Behavioural Ecology of the Komodo Monitor. University of Florida, Gainesville.
AUFFENBERG,W. 1981b Combat behaviour in Varanus bengalensis. J.Bombay N.H.S. 78(1):54-72.
AUFFENBERG,W. 1982.Catch a lizard, use a lizard. Int.Wldf. 12(6):16-19.
AUFFENBERG,W. 1983a. The burrows of Varanus bengalensis. Rec.Zool.Surv.India  80:375-385.
AUFFENBERG,W. 1983b. Courtship behaviour in Varanus bengalensis. In Advances in Herpetology and Evolutionary Biology: Essays in Honor of Ernest E. Williams (Rhodin & Myata eds.): 535-551.
AUFFENBERG,W. 1983c. Notes on feeding behaviour of Varanus bengalensis. J. Bombay N.H.S. 80 (2): 286-302.
AUFFENBERG,W. 1986. The Indian monitor lizard. Sanctuary Asia. 6 (4):327-333.
AUFFENBERG,W. 1988. Gray's Monitor Lizard. University of Florida, Gainesville.
AUFFENBERG,W. 1989. Exploitation of monitor lizards in Pakistan. Unpublished consultancy report to CITES.
AUFFENBERG,W. 1989b. Utilization of monitor lizards in Pakistan. Traffic Bull. 11(1):8-12.
AUFFENBERG,W. 1994. The Bengal Monitor. University Of Florida Press.
AUFFENBERG,W. ARAIN,Q.N. & KURSHID,N. 1991. Mertensiella 2: 7-28. Preferred habitat, home range and movement patterns of Varanus bengalensis in Southern Pakistan.
AUFFENBERG,W. & AUFFENBERG,T. 1990. The reptile tick Aponomma gervaisi as a parasite of monitor lizards in Pakistan and India. Bull. Florida Mus. Nat. Hist. Biol. Sci. 35(1)1-34.
AUFFENBERG,W. & IPE,I.M. 1983. The food and feeding of juvenile Bengal monitors (Varanus bengalensis). J.Bomb.N.H.S. 80:119-124.
AUFFENBERG,W., REHMAN,H., IFFAT,F. & PERVEEN,Z. 1989.  A study of Varanus flavescens.  J.Bombay. Nat.Hist.Soc. 86 (3): 286-307.
AUFFENBERG,W., REHMAN,H., IFFAT,F. & PERVEEN,Z. 1990. Notes on the biology of Varanus griseus koniecznyi. J.Bombay. Nat.Hist.Soc. 87 (1); 26-36.
BAHL,K.N. 1937. Rec. Indian Mus. 39: 133-174. Skull of Varanus monitor.
BAHNOTAR,R.K. & SRIVASTAVA,Y.N. 1985.  Impact of reptiles on food production and environment.  All India Sci. Writers. Soc. Res. Publ. no 2: 116-220, 265-266
BAILON,S. 1992. Escamosos fosiles de facimiento de Moreda (Pliocen medio/superior, Granada, Espana. Revista. espanola de Paleontologia. Numero extraordinario 1992:11-15.
BALSAI,M. 1992. The general care and maintenance of savannah monitors and other popular monitor species. Advanced Vivarium Systems, Lakeside, California.
BANERJEE,V. & BANERJEE,M. 1969. Seasonal variation of erthrocyte number and haemoglobin content in a common Indian lizard Varanus monitor [bengalensis]. Zool.Anz. 182: 203-207.
BARBOUR,T. 1921. Aquatic skinks and arboreal monitors. Copeia 1921 (1): 42-44.
BARBOUR,T. 1943 Defence posture of Varanus gouldii. Copeia 1943(1):56.
BARBOUR,T. & LOVERIDGE,A. 1928.Herpetofauna of Ulugoru and Osambara Mountains, Tanganyika. Bull.Mus.Comp.Zool.Vol L. no.2: 87-265.
BARKER,D.G. 1984. Maintenance and reproduction of Varanus  prasinus kordensis at the Dallas Zoo. Proc. Reptile Breeding Symposium, Columbus:91-92.
BARNETT,B. 1979. Incubation of Varanus gouldii eggs. Herpetofauna 11(1):21-22.
BARNETT,B. 1981. Observations on fish feeding reptiles. Herpetofauna 13 (1): 11-13
BARRETT,C.1928. Lizards in Australian wilds. Bull.New York Zool.Soc. 31(4):
BARRETT,C. 1950. Reptiles of Australia. Cassel, Melbourne.
BARTHOLOMEW,G.A. & TUCKER,V.A. 1964. Size, body temperature, thermal conductance. oxygen consumption and heartrate in Australian Varanid lizards. Physiol.Zool. 37 (4): 341-354.
BARTLETT,R.D. 1981. Notes on captive reproduction of Varanus storri. Bull. Chicago Herp. Soc. 16 (3):65-66.
BARTLETT,R.D. 1982.Initial observations on the captive reproduction of Varanus storri. Herpetofauna 13(2):6-7
BATES,M.F. 1990. Varanus albigularis. Rock Monitor. Hibernation. J.Herpet. Ass. Africa.37:50
BAUER,P. 1972. Cheek to cheek. Int.Wldf. 2(3):54.
BAVERSTOCK,P.R., KING,D., KING,M., BIRRELLJ., KRIEG,M. 1993. The evolution of the species of the Varanidae: microcomplement fixation analysis of serum albumens. Aus.J. Zoology 41(6):621-638.
BAYLESS,M.K. 1994. Zur Fortpflanzungsbiologie des Steppenwaraans (Varanus exanthematicus) Salamandra 30 (2):109-118.
BAYLESS,M.K. & HUFFAKER,R. 1992. Observations of egg deposition and hatching of the svannah monitor (Varanus exanthematicus) in captivity. Varanews 3(1):5-6.
BAYLESS,M.K. & REYNOLDS,T. 1992. Breeding of the savannah monitor lizard in captivity (Varanus exanthematicus). SouthWest. Herp.Soc. 22(1):12-14.
BECKER,H.O., BOHME,W. PERRY,S.F. 1989. Die Lungen morphologie der Warane und ihre systematis-che stammersgeschichichtliche. Bonn. Zool. Beitr. 40 (1):27-56.
BECKER,H.O. 1991. The lung morphology of Varanus yemenensis BOHME et al 1989, and its bearing on the Systematics of Afro-Asian monitor Radiation. Mertensiella 2:29-37.
BEEHLER,B.M. CRILL,W. JEFFERIES,B. & JEFFEREIES,M. New Guinea harpy eagle attempts to capture a monitor lizard. Emu 92(4):246-247.
BEHRMANN,H.J. 1981.Haltung und Nachzucht von Varanus t. timorensis. Salamandra 17(3-4):198-201.
BELCHER,D. 1980. Breeding of Varanus timorensis timorensis at the Rio Grande Zoo. Unpublished Zoo Report
BELLAIRS.A. 1949. Observations on the snout of Varanus and a comparison with that of other lizards and snakes. J.Anat.83:116-146.
BELLAIRS,A. & MILES,A.E.W. 1970. Apparent failure of tooth replacement in monitor lizards. Brit. J. Herp. 1:189-194.
BELS,V.L., GASC,J.P., GOOSSE,V. RENOUS,S. & VERNET,R. 1995. Functional analysis of the throat display in the sand goanna Varanus griseus. J. Zool. Lond. 235:95-116.
BENNETT,A.F. 1972. The effect of activity on oxygen consumption, oxygen debt and heart rate in the lizards Varanus gouldii and Sauromalus hispidus. J. Comp. Physiol. 79:259-280.
BENNETT,A.F. 1973a. Blood physiology and oxygen transport during activity in two lizards. Comp. Biochem. Physiol. 46A:673-690.
BENNETT,A.F. 1973b. Ventilation in two species of lizards during rest and activity. Comp. Biochem. Physiol. 46A:653-671.
BENNETT,D. 1992a.  Reptiles of the Kara Kum. Part 2: The Caspian Monitor. Aquarist and Pondkeeper 56 (11):90-92.
BENNETT,D. 1992b.  A note on the burrows of Varanus griseus caspius around Repetek, Turkmenistan. Journal of the Southwestern Herpetological Society (UK) 2 (1):18-21.
BENNETT,D. 1992c.  A note on Varanus panoptes rubidus (STORR 1980) in Wanjarri, Western Australia. Bull. Brit. Herp. Soc. 39:28-30.
BENNETT,D. 1992d.  Bosc's monitor lizard. Reptilian 1. (2):25-27.
BENNETT,D. 1993a.  A note on a newly hatched Varanus tristis tristis in the Great Victoria Desert, Western Australia. Bull. Brit. Herp. Soc. 44:35-37.
BENNETT,D. 1993b.  A review of some literature concerning the rough-necked monitor lizard Varanus rudicollis. Reptilian 1 (9):7-10.
BENNETT,D. 1994a. Observations on the social interactions of Varanus (Odatria) storri in captivity. Southwestern Herpetological Society Journal (UK) 2(3):2-5.
BENNETT,D. 1994b.  Some remarks on longevity in monitor lizards. Southwestern Herpetological Society Journal (UK) 2(3):27-29.
BENNETT,D. 1995a. Dumeril's monitor lizard (Varanus dumerilii). Reptilian 3 (3):35-37.
BENNETT,D. 1995b. The water monitor, Varanus salvator. Reptilian 3 (8):15-21.
BENNETT,D. In Press.  Some reptiles of the Great Victoria Desert Part III; The Goannas. Aquarist & Pondkeeper 1995/6.
BENNETT,D. In pressb. Notes on foraging behaviour of the water monitor Varanus salvator at Kuala Selangor Nature Park, Selangor, Malaysia. Southwestern Herpetological Society Journal (UK).
BENNETT,D. & AKONNOR,W.O. 1995. A preliminary investigation into the ecology of the monitor lizard Varanus exanthematicus in the coastal savannah of Ghana. Unpublished report to the Department of Game and Wildlife, Accra, Ghana. March 1995. 17pp.
BENNETT,D. & AKONNOR,W.O. ms. The ecology and exploitation of  monitor lizards  in the coastal savannah of Ghana.
BENNETT,D. & LIM,B,L. In press. A note on the distribution of Varanus rudicollis and Varanus dumerilii in Peninsular Malaysia. Malay.Nat.J.
BENNETT,D. & SHIMANSKAYA,M. 1991.  A review of some Soviet literature concerning the grey monitor Varanus griseus caspius (EICHWALD). Varanews 1(8):5-8.
BERANEK,S. 1985. Varanus bengalensis (in Czeckoslovakian). Ark.Ter. 28(4):28.
BERG,A. 1913. Unsur Buurtwaran (Varanus varius). Blat. Aquar.Terrar. Kunde 24:717-719.
BERNEY,F.L. 1936. Gould's monitor. Queensland Nat. 10(1)
BHANOTAR,R.K. & SRIVASTAVA,Y.N. 1985. Impact of reptiles on food production and environment. All Ind.Sci.Writers. Soc. (AISWS) Res.Publ. 1985 (2):116-120.
BHARATAN,E. 1971.Monitor preying on a cobra. J.Bombay Nat.Hist.Soc. 68(3):838 .
BICKLER,P.E. & ANDERSON,R.A. 1986. Ventilation, gas exchange, and aerobic scope in a small monitor lizard, Varanus gilleni. Physiol.Zool. 59(1):76-83.
BIEBL,H. 1993. Erfolgreiche Nachzucht von Varanus prasinus prasinus. Monitor (Frankfurt) 2(2):33.
BIEBL,H. 1994. Erfolgreiche Nachzucht von Beccar's Smaragdwaran Varanus prasinus beccarii. Monitor (Frankfurt) 3(1):37-40.#
BIEBL,H. 1995. Haltungsrekord bei der pflege eines Dumeril-Warans (Varanus dumerilii) nebst eininger personlichen Anmerkungen zur Haltung und Pflege dieses speziellen Examplares. Monitor (Frankfurt) 4(1):24-29.
BILLIAU,F. 1972. Voedselweigering en genezing van een woestijnvaraan. Lacerta 31 no2:30-32.
BIRCHARD,G.F. ,WALSH.T.& REIBER,C.L.1995. Oxygen uptake by Varanus komodoensis eggs. Physiol.Zool. 68(4):622-
BISHAI,H.M. 1959. The anatomy and histology of the alimentary tract of the lizard Varanus griseus Daud. Bull. Fac.Science 35:53-69. (Reprinted 1960. Cairo University Press).
BISWAS,S. & ACHARJYO,L.N. 1977. Notes on the ecology and biology of some reptiles of Nadankanan Biological Park, Orissa. Rec.Zool.Surv.India 73:95-109.
BISWAS,K. & KARR,S. 1981. Some observations on nesting habits and biology of Varanus  salvator in Bhitarkanika Sanctuary, Orissa. J.Bombay.Nat.Hist.Soc. 78(2):303-308.
BLAIR,L. & BLAIR,L. 1988. Ring of Fire. Pgs 191-211. Bantam Press, London.
BOHME,W. 1988. Der Arguswaran (Varanus panoptes STORR, 1980) auf Neuguinea. Varanus panoptes horni. ssp.n. Salamandra 24 (2\3):87-101. #
BOHME,W. 1988. Zur Genitalmorphologie der Sauria: funktionelle und stammesgeschichtliche Aspekte. Bonn Zool.Monog. 27.
BOHME,W. 1991. The identity of Varanus gouldii (GRAY 1838), and the nomenclature of the Varanus gouldii species complex. Mertensiella 2:38-41.
BOHME,W. 1991. New findings on the hemipenal morphology of monitor lizards and their systematic implications. Mertensiella 2: 42-49.
BOHME,W, HORN,H.G. & ZIEGLER,T. 1994. Zur taxonomie der Pazifikwarane (Varanus indicus Komplex): Revalidierung von Varanus doreanus (A.B. MEYER, 1874) mit Beschreibung einer neuen Unterart. Salamandra 30(2):119-142.#
BOHME,W., FRITZ,J.P. & SCHUTTE,F. 1987. Neuentdeckung einer Grossechse (Varanus) aus der Arabischen Republik Jemen. Herpetofauna 9 (46):13-20. #
BOHME,W., U.JOGER & B.SCHATTI. 1989. A new monitor lizard from Yemen, with notes on ecology, phylogeny and zoogeograophy. Fauna of Saudi Arabia 10:433-448.
BOM,A. & BOM,I. 1989. De steppenvaraan (Varanus exanthematicus microstictus) in Reptilienzoo Iguana.  Lacerta 48(1):2-5.
BONE,W.H. 1920. The booming lizard of Australia.Science 52:273.
BOONRATANA,R. 1988. Distributional Survey and trade of Varanus species in Thailand. Unpublished consultancy report to CITES..
BOONRATANA,R. 1988. Additional report on trade of Varanus spp. in Thailand (Supplement to above).
BOONSONG,L. 1969. Monitors (Varanus) of Thailand. Conservation News S.E. Asia. 8 (DEC):31-32.
BORODIN,A.M. ET AL. 1978. U.S.S.R. Red Data Book. Lesnaya Promyshlenn ost. Moscow. (in Russian)
BOSCH,H. 1991. The pentastome Elenia HEYMONS 1932 a parasite genus specific to monitors from the Australian region - Biology and Aspects of Host Specificity. Mertensiella 2:50-56.
BOULENGER,G.A. 1885. Catalogue of lizards in the British Museum (Natural History) 2.
BOURQUIN,O. 1991. Varanus niloticus reproduction. J.Herpet.Ass.Africa 39:21.
BOURQUIN,O. & CHANNING,A. 1980. Herpetofauna of the Natal Drakensburg: an annotated checklist. Lammergeyer 30:1-20.
BOWERS,M.J. 1981. Varanus salvator bred in captivity. San Antonio News from the Zoo. 7(2):1-2.
BOWKER,R.G. 1984. Precision of thermoregulation of some African lizards. Physiol. Zool. 57 (4): 401-412.
BOYCOTT,R.C. & MORGAN,D.R. 1988. Reproductive and growth data of some South African lizards. Herpet. Ass. Afr. 35:15-18.
BOYER,D.M. & LAMOREAUX,W.E. 1983. Captive reproduction in Varanus gilleni at the Dallas Zoo. Proc.Rept.Symp.Capt.Prop.Husb. 84(7):59-63.
BOYLAN ,T. 1995. Field observations, captive breeding and growth rates of the lace monitor Varanus varius. Herpetofauna (Sydney) 25 (1):10-14.
BRANCH,W.R. 1982. Hemipeneal morphology of platynotan lizards.. J. Herpetology. 16:16-38.
BRANCH,W.R. 1988. Varanus exanthematicus albigularis : rock leguaan: egg size. J. Herpet.Assoc. Africa. 35:39.
BRANCH,W.R. 1988.  Bill Branch's field guide to the snakes and other reptiles of southern Africa. New Holland, London.
BRANCH,W.R. 1989. Alternative life-history styles in reptiles. In Bruton,M.N. (ed). Alternative life-history styles of animals. Kluwer Academic Publishers, Dordrecht. pp. 127-151.
BRANCH,W.R. 1991. The Regenia Registers of `Gogga' Brown (1869-1909) "Memoranda on a species of monitor or varan". Mertensiella 2:57-110.
BRANCH,W.R. & BRAACK,H.H. 1987. Reptiles and amphibians of the Addo Elephant National Park. Koedoe 30:61-11.
BRANCH,W.R. & BRANCH.T.C. 1992. Varanus niloticus niloticus. Water monitor. Size. J.Herpet.Ass.Africa 41:41.
BRANCH,W.R. & ERASMUS,H. 1982. Notes on reproduction in the South African water monitor Varanus niloticus niloticus. J.Herpet.Soc.Afr. 28:4-9.
BRANDENBURG,T. 1983. Monitors in the Indo-Australian Archipelago. 1-121. Thesis, University of Leiden.
BRATTSTROM,B.H. 1973. Rate of heat loss by large Australian monitor lizards. Bull. South Calif. Acad. Sci. 72: 52-54.
BRAYSHER,M. & GREEN,B. 1970. Absorption of water and electrolytes from the cloaca of an Australian lizard Varanus gouldii (Gray). Comp. Biochem. Physiol. 35:607-614.
BREDL,J. 1987. First captive breeding of the perentie (Varanus giganteus). Thylacinus 12(1):2-3.
BREDL,J. & HORN,H.G. 1987. Uber die Nachzucht des australischen Riesenwarens Varanus giganteus. Salamandra 23(2/3):90-96.
BREDL,J. & SCHWANER,T.D. 1983. First record of captive propogation of the lace monitor (Varanus varius). Herpetofauna 15(1):20-21.
BROADBENT,K. 1910.  Birds of Cardwell and Herbert River Districts. Emu 10:233-245
BROADLEY,D.G. 1971. Reptiles and amphibians of Zanzibar. The Puku 6:1-143.
BROER,W. & HORN,H.G. 1985. Erfahrungen bei Verwendung eines Motorbruters zur Zeitigung von Reptilieneiern. Salamandra 21(4):304-310.
BRONGERSMA,L.D. 1932. Uber die Eiablage und die Eier von Varanus komodoensis. Zool. Garten 5:45-48.
BRONGERSMA,L.D. 1947. Zoological notes from Port Dickson, I. Amphibians and reptiles. Zoologische Mededelingen, Rijksmuseum van Natuurlijke Historie. 27:300-308.
BRONGERSMA,L.D. 1958. On an extinct species of Varanus from the island of Flores. Zool.Meded.Rijks Mus.Nat.Hist.Leiden. 36 (7):114-125.
BROOM,R. 1898 Proc.Linn.Soc.New South Wales 22:639-645.On the lizards of the Chillagoe District, northern Queensland.
BROOKER,M.G. & WOMBEY,J.C. 1978. Some notes on the herpetofauna of the western Nullabar plain, Western Australia. West Aus. Nat. 14 (2): 36-41.
BROOKER,M.G. & WOMBEY,J.C. 1986. Some observations on the herpetofauna of the Macquarie Marshes region, N.S.W., with special reference to Chelidae. Aust. Zool. 23(1):1-4.
BROTZLER,A. 1965. Freund Kolner Zoo. 8(3):89. Mertens-Wasserwarane (Varanus mertensi Glauert 1951) zuchteten in der Wilhelma.
BROUGHTON,M. 1936. Nat. Geog. Mag. 70:321-331. A modern dragon hunt on Komodo.
BROWN,K.W. 1983. The green tree monitor Varanus prasinus. Scientific Committee meeting report, Chester Zoo.
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About Mampam
Bye Bye Butaan


Butaan start to visit fruiting trees before they are large enough to swallow the fruits. They make repeat journeys to trees, perhaps to reinforce memory of the position of the tree. If the youngster survives it may continue to use this tree for many decades. Fruiting trees like this are a vital resource for entire populations of butaan. Learn more >

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The Butaan Project
The Butaan Project


Since 1999 the Butaan Project has been studying the rare, endangered, and unique fruit-eating monitor lizards of the Philippines.  Butaan is just one of several races of frugivorous monitor lizards in the Philippines ("Pandan Biawak"), all of which are of at least as great a conservation concern as the Komodo dragon, but receive virtually none of the attention. Pandan Biawak occur only in lowland dipterocarp forest. The first species (Butaan) was discovered in 1845 and not seen alive by a scientist until the late 1970s. The next species (Mabitang) was discovered in 2001. Other species remain undescribed, and some may have gone extinct without ever having been recognised.





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